Checkpoint Proteins Bub1 and Bub3 Delay Anaphase Onset in Response to Low Tension Independent of Microtubule-Kinetochore Detachment

？ Taxol-sensitive yeast model allows separation of kinetochore tension and attachment ？ Low tension at attached kinetochores generates a delay in anaphase onset ？ The tension-specific delay mechanism requires the SAC proteins Bub1 and Bub3 ？ The tension mechanism does not require the core SAC proteins Mad1, Mad2, and Mad